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Brood discrimination, nest mate discrimination, and determinants of social behavior in facultatively quasisocial beetles (Nicrophorus spp.). Behavioral Ecology. 4(4):332-339.. 1993.
Burying beetles: intraspecific interactions and reproductive success in the field. Ecological Entomology. 9:195-203.. 1984.
The effect of population structure on the evolution of mutualism: a field test involving burying beetles and their phoretic mites. The American Naturalist. 121:851-870.. 1983.
Food discrimination and ovarian development in burying beetles (Coleoptera: Silphidae: Nicrophorus). Annals of the Entomological Society of America. 77:165-170.. 1984.
Genetic polymorphism for carrier preference in a phoretic mite. Annals of the Entomological Society of America. 75:293-296.. 1982.
Prudent predation: a field study involving three species of tiger beetles. Oikos. 31:128-136.. 1978.
Species packing and temperature dependent competition among burying beetles (Silphidae, Nicrophorus). Ecological Entomology. 9:205-216.. 1984.